Bryophyta and Tracheophyta

Bryophytes include the mosses, liverworts, and hornworts. Bryophytes are the simplest of plants (excluding the algae, which are not considered plants by most botanists). Bryophytes are small, seldom exceeding 6-8 in (15-20 cm) in height, and usually much smaller. They are attached to the substrate (ground, rock, or bark) by rhizoids, which are one or a few-celled, root-like threads that serve only for anchoring and are not capable of absorbing water and nutrients from the substrate. Brypohytes lack vascular tissue (the specialized cells grouped together to pipe water and nutrients to various parts of the body), or in the rare cases when this tissue is present, it is not well differentiated. The leaves of bryophytes are technically not true leaves, because in most species they lack vascular tissue. However, they are functionally equivalent to leaves, containing chlorophylls a and b for photosynthesis. Leaves are usually one-cell thick, except for the midrib, which may be up to 15 cells thick. Bryophytes satisfy their nutritional requirements by absorbing minerals from dust, rainfall, and water running over their surface.

The life cycle of bryophytes is characterized by an alternation of generations, one of which is a multicellular, diploid individual called a sporophyte, having two of each type of chromosome per cell. This stage alternates with multicellular, haploid individual called the gametophyte, with only one of each type of chromosome per cell, as is also the case with animal sperm. Bryophytes are unique among plants in that the dominant, conspicuous generation is the haploid gametophyte. In all other plants, the dominant stage is the diploid sporophyte.

Most reproduction of bryophytes is asexual, occurring by fragmentation of body parts, and by the production of specialized vegetative units called gemmae. Gemmae may be produced as microscopic plates (in the genus Tetraphis), as bulbils in the axils of leaves (in Pohlia), or as microscopic filaments (in Ulota). When sexual reproduction occurs, it always involves a flagellated sperm (produced in a specialized organ called an antheridium) that must swim through water to reach an egg located in a specialized, flask-shaped organ (the archegonium). The antheridia and archegonia are surrounded by a layer of sterile cells, which protects the sex organs from mechanical damage and desiccation.